A mother and calf muskox along the coastline of Alaska's Arctic National Wildlife Refuge. July, 2010.
A male Ruffed Grouse thumps its wings as part of a breeding display to attract female grouse. Kluane, Yukon. May, 2013.
A Great Blue Heron on a sandstone shoreline, Puerto Los Gatos, Baja California.
An African Eagle Owl seeks shade from the unrelenting heat of the Kalahari Desert. South Africa. December, 2011.
A bumblebee searches for nectar in fireweed flowers. Whitehorse, Yukon. July, 2011.
A Great Grey Owl flies towards the forest with a vole it has just caught. Kluane, Yukon. August, 2013.
A lion lounges in brush in the Kalahari Desert, in northwestern South Africa. December, 2011.
The shadow of a new lighthouse on the bricks of the old lighthouse. Isla San Jose, Baja California. November, 2013.
A Long-tailed Widowbird shakes out its tail. Malolotja Preserve, Swaziland. January, 2012.
A Pacific Loon floats in a tundra pond, near the Turner River in the Arctic National Wildlife Refuge. July, 2010.
Snowy Egrets squabble above a fishing pier on Sanibel Island, Florida. February, 2008.
A Dall Sheep flees a Grizzly Bear on the flanks of Sheep Mountain, Yukon. May, 2012.
A Double-collared Sunbird bathes in water trapped in an aloe leaf, in Swaziland's Malolotja Preserve.
A springbok "pronks" in the Kalahari Desert, in northwestern South Africa. December, 2011.
Clouds tatter along rocky crags of the Mt Vulcan massif, in the Yukon's Kluane Mountains. June, 2013.
Storm clouds approach the nest of a Pacific Loon, near the Turner River in the Arctic National Wildlife Refuge. July, 2010.
A Jackass Penguin shakes itself out. Cape of Good Hope peninsula, South Africa. December, 2013.
Mist shrouds an island in Kluane Lake. June, 2013.
Great Blue HeronBaja California